141-Year-Old Galapagos Tortoise "Gramma" Dies at San Diego Zoo: A Century-Long Legacy of Conservation

Nov 25, 2025 05:50 pm CST

The San Diego Zoo mourns the loss of Gramma, a 141-year-old Galapagos tortoise who lived through two World Wars and 20 US presidencies. Known as the "Queen of the Zoo," Gramma's remarkable life highlights the conservation challenges facing these endangered reptiles, while recent breeding successes offer hope for the species' survival.